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The Welsh camp has been blighted with injuries since the announcement of the squad just over a week ago.
Welsh captain Ryan Jones will miss the first test against Australia on November 6 with a calf tear with the captaincy passing over to Scarlets skipper Matthew Rees.
Full-back Lee Byrne and wing Leight Halfpenny have both been ruled out for the duration of the November internationals, Byrne fractured his third metacarpal in his right hand at the weekend and Halfpenney has damaged ankle ligaments.
Dragons winger Aled Brew and Blues full-back Chris Czekaj have both been added to squad as cover.
Scarlets back row Rob McCusker has also withdrawn from the squad in order to undergo a hernia operation.
McCusker faces a six week lay-off after the operation and will be targeting a return to action in time for the Scarlets' back-to-back fixtures against Benetton Treviso in the Heineken Cup in December.
Uncapped Scarlets winger George North is still recovering from a fracture to his fibula but is expected to play some part in the series.
Wales Squad:
Gethin JENKINS (CARDIFF BLUES), Paul JAMES (OSPREYS), Adam Rhys JONES (OSPREYS), John YAPP (CARDIFF BLUES), Scott ANDREWS (CARDIFF BLUES), Matthew REES (SCARLETS), Huw BENNETT (OSPREYS), Richard HIBBARD (OSPREYS), Alun Wyn JONES (OSPREYS), Bradley DAVIES (CARDIFF BLUES), Deiniol JONES (CARDIFF BLUES), Ian GOUGH (OSPREYS), Dan LYDIATE (NEWPORT GWENT DRAGONS), Jonathan THOMAS (OSPREYS), Ryan JONES (OSPREYS), Sam WARBURTON (CARDIFF BLUES), Martyn WILLIAMS (CARDIFF BLUES), Rob McCUSKER (SCARLETS), Michael PHILLIPS (OSPREYS), Richard REES (CARDIFF BLUES), Tavis KNOYLE (SCARLETS), Stephen JONES (SCARLETS), Dan BIGGAR (OSPREYS), James HOOK (OSPREYS), Andrew BISHOP (OSPREYS), Jonathan DAVIES (SCARLETS), Tom SHANKLIN (CARDIFF BLUES), Shane WILLIAMS (OSPREYS), Leigh HALFPENNY (CARDIFF BLUES), Will HARRIES (NEWPORT GWENT DRAGONS), George NORTH (SCARLETS), Lee BYRNE (OSPREYS), Tom PRYDIE (OSPREYS).
Scotland:
Gloucester flanker Alasdair Strokosch has withdrawn from the squad after damaging knee ligaments in his side's victory over Harlequins at the weekend.
The 27-year-old is expected to be sidelined for six weeks, ruling him out of the autumn series and will have a race against time on his hands to be fit for Gloucester's trip to La Rochelle on December 12 in the Amlin Challenge Cup.
Andy Robinson name four uncapped players in his squad and they are, 22-year-old Glasgow Warriors stand-off Ruaridh Jackson, Edinburgh duo, scrum-half Greig Laidlaw and full-back/wing Jim Thompson along with Northampton Saints utility back Joe Ansbro.
Scotland squad:
Backs - Joe Ansbro (Northampton Saints), Mike Blair (Edinburgh), Ben Cairns (Edinburgh), Max Evans (Glasgow Warriors), Alex Grove (Edinburgh), Ruaridh Jackson (Glasgow Warriors), Greig Laidlaw (Edinburgh), Rory Lamont (Toulon), Sean Lamont (Scarlets), Rory Lawson (Gloucester), Graeme Morrison (Glasgow Warriors), Dan Parks (Cardiff Blues), Chris Paterson (Edinburgh), Hugo Southwell (Stade Francais), James Thompson (Edinburgh), Nikki Walker (Ospreys)
Forwards - John Barclay (Glasgow Warriors), Kelly Brown (Saracens), Geoff Cross (Edinburgh), Alasdair Dickinson (Gloucester), Ross Ford (Edinburgh), Richie Gray (Glasgow Warriors), Dougie Hall (Glasgow Warriors), Jim Hamilton (Gloucester), Nathan Hines (Leinster), Allan Jacobsen (Edinburgh), Scott Lawson (Gloucester), Moray Low (Glasgow Warriors), Alan MacDonald (Edinburgh), Scott MacLeod (Edinburgh), Euan Murray (Northampton Saints), Ross Rennie (Edinburgh), Alasdair Strokosch (Gloucester) and Richie Vernon (Glasgow Warriors).
Not considered due to injury - Johnnie Beattie (Glasgow Warriors), Chris Cusiter (Glasgow Warriors), Simon Danielli (Ulster), Nick De Luca (Edinburgh), Phil Godman (Edinburgh) and Alastair Kellock (Glasgow Warriors
Ireland:
Double Heineken Cup winner Marcus Horan hs been ruled out for several weeks with a hand injury and uncapped Connacht prop Brett Wilkinson has been called into the squad as cover.
In positive news Leinster's Luke Fitzgerald has been added to the squad after recovering from injury.
The British Lion tourist was not included in the original 34-man squad, but has recovered sufficiently to join his team mates in the Ireland camp based in Limerick.
Ireland Squad:
Forwards (19) -Rory Best (Banbridge/Ulster) Tony Buckley (Shannon/Munster) Tom Court (Malone/Ulster) Sean Cronin (Buccaneers/Connacht) Stephen Ferris (Dungannon/Ulster) John Hayes (Bruff/Munster) Cian Healy (Clontarf/Leinster) Jamie Heaslip (Naas/Leinster) Marcus Horan (Shannon/Munster) Denis Leamy (Cork Constitution/Munster) John Muldoon (Galwegians/Connacht) Donncha O'Callaghan (Cork Constitution/Munster) Sean O'Brien (Clontarf/Leinster) Mick O'Driscoll (Cork Constitution/Munster) Mike Ross (Clontarf/Leinster) Donnacha Ryan (Shannon/Munster) Devin Toner (Lansdowne/Leinster) * Damien Varley (Garryowen/Munster) David Wallace (Garryowen/Munster)
Backs (15) -Isaac Boss (Terenure College/Leinster) Tommy Bowe (Ospreys) Gordon D'Arcy (Lansdowne/Leinster) Gavin Duffy (Galwegians/Connacht) Keith Earls (Thomond/Munster) Shane Horgan (Boyne/Leinster) Rob Kearney (UCD/Leinster) Johne Murphy (Young Munster/Munster) * Ronan O'Gara (Cork Constitution/Munster) Brian O'Driscoll (UCD/Leinster) Eoin Reddan (Lansdowne/Leinster) Jonathan Sexton (St. Mary's College/Leinster) Peter Stringer (Shannon/Munster) Andrew Trimble (Ballymena/Ulster) Paddy Wallace (Ballymena/Ulster)
England:
Martin Johnson has a number of injury concerns and has called four players up to his Elite Player Squad as cover.
Sale Sharks' points machine Charlie Hodgson arrives with Toulon's Jonny Wilkinson suffering with a shoulder injury.
Harlequins prop Joe Marler comes in for Bath's David Wilson who has a sore back. Northampton Saints flanker Tom Wood has been include, with Hendre Fourie suffering from brusied ribs.
Simon Shaw is suffering with a calf injury so club-mate Dan Ward-Smith joins the camp at Penny Hill park.
England Squad:
Forwards: Dan Cole, Paul Doran-Jones, Tim Payne, Andrew Sheridan, David Wilson, George Chuter, Dylan Hartley, Steve Thompson, Dave Attwood, Courtney Lawes, Tom Palmer, Simon Shaw, Tom Croft, Hendre Fourie, James Haskell, Lewis Moody, Joe Worsley, Nick Easter.
Backs: Danny Care, Ben Youngs, Toby Flood, Jonny Wilkinson, Riki Flutey, Shontayne Hape, Mike Tindall, Dominic Waldouck, Chris Ashton, Matt Banahan, Mark Cueto, David Strettle, Delon Armitage, Ben Foden.
Italy:
Italy are once again lead by inspiration No8 Sergio Parisse and havent suffered any major set-backs since Nick Mallet announced his squad.
The squad includes two-uncapped players including Benetton Treviso star Tommaso Benvenuti, who has scored four tries in two Heineken Cup appearances so far this season.
Italy training squad:
Forwards: Robert Barbieri (Treviso, 7 caps), Mauro Bergamasco (Stade Français, 84 caps), Valerio Bernabo' (Treviso, 15 caps), Lorenzo Cittadini (Treviso, 3 caps), Martin Castrogiovanni (Leicester , 68 caps), Carlo Antonio Del Fava (Aironi, 45 caps), Santiago Dellape' (Racing Metro , 57 caps), Paul Derbyshire (Treviso, 5 caps), Quintin Geldenhuys (Aironi , 13 caps), Leonardo Ghiraldini (Treviso, 31 caps), Andrea Lo cicero (Racing Metro, 78 caps), Fabio Ongaro (Aironi, 71 caps), Sergio Parisse(C) (Stade Français, 69 caps), Antonio Pavanello (Treviso, 5 caps), Salvatore Perugini (Aironi , 73 caps), Alessandro Zanni (Treviso, 44 caps).
Backs:Tommaso Benvenuti (Treviso), Mirco Bergamasco (Racing Metro, 73 caps), Riccardo Bocchino (Aironi , 4 caps), Kristhoper Burton (Treviso, 4 caps), Gonzalo Canale (Clermont, 58 caps), Pablo Canavosio (Aironi , 30 caps), Denis Majstorovic (Estra I Cavalieri Prato ), Andrea Masi (Racing Metro , 52 caps), Luke Mclean (Treviso, 21 caps), Luciano Orquera (Brive, 14 caps), Michele Sepe (Treviso, 3 caps), Alberto Sgarbi (Treviso, 5 caps), Tito Tebaldi (Aironi, 13 caps), Giulio Toniolatti (Aironi, 5 caps),
